An endemic rat species complex is evidence of moderate environmental changes in the terrestrial biodiversity centre of China through the late Quaternary.

A slight trend of recent population decline was identified in the lineage of Yunnan/SE Tibet in N. andersoni ( Fig. 4F ).

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excelsior expanded with a clear trend of westward extension to the edge of the southwestern QHTP- Himalaya region during the LGM (~0.025 Mya).

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