Mean (±se; t-test) net displacements were 5 m eastwards (± 23 m; t = 0.22, df = 5, P = 0.84) and 6 m southwards (± 13 m; t = -0.50, df = 5, P = 0.64) for the flight-experienced beetles, and 76 m westwards (± 50 m; t = -1.51, df = 3, P = 0.23) and 114 m southwards (± 34 m; t = -3.33, df = 3, P = 0.045) for the flight-naïve beetles ( Fig 4 ), indicating that southward directionality, or drift, was only marginally significant for the flight-naïve beetles.
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