Selection at a genomic region of major effect is responsible for evolution of complex life histories in anadromous steelhead.

All six sub-basins showed a trend between spawning tributary arrival date and genotype class, with five out of the six having statistically different arrival week means between “premature” and “mature” genotypes (Fig. 4 , Additional file 1 : Figure S3 and Table S5).

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This region also includes a highly significant intergenic region between greb1L and rock1 which may play a regulatory role in expression of these genes.

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