approaching significanceP =0.0439
Results showed that mtDNA F ST differentiation was positively related to geographic distance and negatively related to PC1 (MMRR R 2 =0.322, P <0.001), while nuDNA F ST was negatively associated with precipitation differences (considering synergy of multiple precipitation variables), approaching significance (MMRR R 2 =0.109, P =0.0439).