Temperature and land use change are associated with <i>Rana temporaria</i> reproductive success and phenology.

Hatch date Site T04 (Upper Teesdale, North Pennines) exhibited a negative trend over time in hatch date, indicative of earlier hatching every year ( p = −0.4, p.adj = 0.045; Fig. 3A ).

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Reproductive success Though several sites showed increases over time in the percentage of dead spawn, only site T11 (Yr Wyddfa, Wales) had a significant trend when correcting for multiple testing (cor = 0.54, p.adj = 0.02; Fig. 4A ).

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